649 research outputs found

    A users manual for a computer program which calculates time optical geocentric transfers using solar or nuclear electric and high thrust propulsion

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    This manual is a guide for using a computer program which calculates time optimal trajectories for high-and low-thrust geocentric transfers. Either SEP or NEP may be assumed and a one or two impulse, fixed total delta V, initial high thrust phase may be included. Also a single impulse of specified delta V may be included after the low thrust state. The low thrust phase utilizes equinoctial orbital elements to avoid the classical singularities and Kryloff-Boguliuboff averaging to help insure more rapid computation time. The program is written in FORTRAN 4 in double precision for use on an IBM 360 computer. The manual includes a description of the problem treated, input/output information, examples of runs, and source code listings

    Coexistence of competitors mediated by nonlinear noise

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    Stochastic reaction-diffusion equations are a popular modelling approach for studying interacting populations in a heterogeneous environment under the influence of environmental fluctuations. Although the theoretical basis of alternative models such as Fokker- Planck diffusion is not less convincing, movement of populations is most commonly modelled using the diffusion law due to Fick. An interesting feature of Fokker-Planck diffusion is the fact that for spatially varying diffusion coefficients the stationary solution is not a homogeneous distribution – in contrast to Fick’s law of diffusion. Instead, concentration accumulates in regions of low diffusivity and tends to lower levels for areas of high diffusivity. Thus, we may interpret the stationary distribution of the Fokker-Planck diffusion as a reflection of different levels of habitat quality. Moreover, the most common model for environmental fluctuations, linear multiplicative noise, is based on the assumption that individuals respond independently to stochastic environmental fluctuations. For large population densities the assumption of independence is debatable and the model further implies that noise intensities can increase to arbitrarily high levels. Therefore, instead of the commonly used linear multiplicative noise model, we implement environmental variability by an alternative nonlinear noise term which never exceeds a certain maximum noise intensity. With Fokker-Planck diffusion and the nonlinear noise model replacing the classical approaches we investigate a simple invasive system based on the Lotka-Volterra competition model. We observe that the heterogeneous stationary distribution generated by Fokker-Planck diffusion generally facilitates the formation of segregated habitats of resident and invader. However, this segregation can be broken by nonlinear noise leading to coexistence of resident and invader across the whole spatial domain, an effect that would not be possible in the non-spatial version of the competition model for the parameters considered here

    Robust ecological pattern formation induced by demographic noise

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    We demonstrate that demographic noise can induce persistent spatial pattern formation and temporal oscillations in the Levin-Segel predator-prey model for plankton-herbivore population dynamics. Although the model exhibits a Turing instability in mean field theory, demographic noise greatly enlarges the region of parameter space where pattern formation occurs. To distinguish between patterns generated by fluctuations and those present at the mean field level in real ecosystems, we calculate the power spectrum in the noise-driven case and predict the presence of fat tails not present in the mean field case. These results may account for the prevalence of large-scale ecological patterns, beyond that expected from traditional non-stochastic approaches.Comment: Revised version. Supporting simulation at: http://guava.physics.uiuc.edu/~tom/Netlogo

    Dynamical density functional theory for the dewetting of evaporating thin films of nanoparticle suspensions exhibiting pattern formation

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    Recent experiments have shown that the striking structure formation in dewetting films of evaporating colloidal nanoparticle suspensions occurs in an ultrathin `postcursor' layer that is left behind by a mesoscopic dewetting front. Various phase change and transport processes occur in the postcursor layer, that may lead to nanoparticle deposits in the form of labyrinthine, network or strongly branched `finger' structures. We develop a versatile dynamical density functional theory to model this system which captures all these structures and may be employed to investigate the influence of evaporation/condensation, nanoparticle transport and solute transport in a differentiated way. We highlight, in particular, the influence of the subtle interplay of decomposition in the layer and contact line motion on the observed particle-induced transverse instability of the dewetting front.Comment: 5 pages, 5 figure

    Nonequilibrium plankton community structures in an ecohydrodynamic model system

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    International audienceDue to the local and global impacts of algae blooms and patchiness on water quality, carbon cycling and climate, models of plankton dynamics are of current interest. In this paper, the temporal and spatial patterns in natural plankton communities are interpreted as transient and stationary nonequilibrium solutions of dynamical nonlinear interaction-diffusion-advection systems. A simple model of phytoplankton-zooplankton dynamics (Scheffer, 1991) is presented in space and time. After summarizing the local properties as multiple stability and oscillations, the emergence of spatial and spatio- temporal patterns is considered, accounting also for diffusion and weak advection. In order to study the emergence and stability of these structures under hydrodynamic forcing, the interaction- diffusion-advection model is coupled to the hydrodynamic equations. It is shown, that the formation of nonequilibrium spatio-temporal density patterns due to the interplay of the deterministic nonlinear biological interactions and physical processes is a rare occurrence in rapidly flowing waters. The two-timing perturbation technique is applied to problems with very rapid single-directed steady flows. A channel under tidal forcing serves as and example for a system with a relatively high detention time of matter. Generally, due to the different time and length scales of planktic interactions, diffusion and transport, initial nonequilibrium plankton patches are simply moved through the system unless the strong hydrodynamic forces do not destroy them before

    Ca(2+ )regulation in the absence of the iplA gene product in Dictyostelium discoideum

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    BACKGROUND: Stimulation of Dictyostelium discoideum with cAMP evokes an elevation of the cytosolic free Ca(2+ )concentration ([Ca(2+)](i)). The [Ca(2+)](i)-change is composed of liberation of stored Ca(2+ )and extracellular Ca(2+)-entry. The significance of the [Ca(2+)](i)-transient for chemotaxis is under debate. Abolition of chemotactic orientation and migration by Ca(2+)-buffers in the cytosol indicates that a [Ca(2+)](i)-increase is required for chemotaxis. Yet, the iplA(- )mutant disrupted in a gene bearing similarity to IP(3)-receptors of higher eukaryotes aggregates despite the absence of a cAMP-induced [Ca(2+)](i)-transient which favours the view that [Ca(2+)](i)-changes are insignificant for chemotaxis. RESULTS: We investigated Ca(2+)-fluxes and the effect of their disturbance on chemotaxis and development of iplA(- )cells. Differentiation was altered as compared to wild type amoebae and sensitive towards manipulation of the level of stored Ca(2+). Chemotaxis was impaired when [Ca(2+)](i)-transients were suppressed by the presence of a Ca(2+)-chelator in the cytosol of the cells. Analysis of ion fluxes revealed that capacitative Ca(2+)-entry was fully operative in the mutant. In suspensions of intact and permeabilized cells cAMP elicited extracellular Ca(2+)-influx and liberation of stored Ca(2+), respectively, yet to a lesser extent than in wild type. In suspensions of partially purified storage vesicles ATP-induced Ca(2+)-uptake and Ca(2+)-release activated by fatty acids or Ca(2+)-ATPase inhibitors were similar to wild type. Mn(2+)-quenching of fura2 fluorescence allows to study Ca(2+)-influx indirectly and revealed that the responsiveness of mutant cells was shifted to higher concentrations: roughly 100 times more Mn(2+ )was necessary to observe agonist-induced Mn(2+)-influx. cAMP evoked a [Ca(2+)](i)-elevation when stores were strongly loaded with Ca(2+), again with a similar shift in sensitivity in the mutant. In addition, basal [Ca(2+)](i )was significantly lower in iplA(- )than in wild type amoebae. CONCLUSION: These results support the view that [Ca(2+)](i)-transients are essential for chemotaxis and differentiation. Moreover, capacitative and agonist-activated ion fluxes are regulated by separate pathways that are mediated either by two types of channels in the plasma membrane or by distinct mechanisms coupling Ca(2+)-release from stores to Ca(2+)-entry in Dictyostelium. The iplA(- )strain retains the capacitative Ca(2+)-entry pathway and an impaired agonist-activated pathway that operates with reduced efficiency or at higher ionic pressure

    Taxis-driven pattern formation in a predator-prey model with group defense

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    We consider a reaction-diffusion(-taxis) predator-prey system with group defense in the prey. Taxis-driven instability can occur if the group defense influences the taxis rate (Wang et al., 2017). We elaborate that this mechanism is indeed possible but biologically unlikely to be responsible for pattern formation in such a system. Conversely, we show that patterns in excitable media such as spatiotemporal Sierpinski gasket patterns occur in the reaction-diffusion model as well as in the reaction-diffusion-taxis model. If group defense leads to a dome-shaped functional response, these patterns can have a rescue effect on the predator population in an invasion scenario. Preytaxis with prey repulsion at high prey densities can intensify this mechanism leading to taxis-induced persistence. In particular, taxis can increase parameter regimes of successful invasions and decrease minimum introduction areas necessary for a successful invasion. Last, we consider the mean period of the irregular oscillations. As a result of the underlying mechanism of the patterns, this period is two orders of magnitude smaller than the period in the nonspatial system. Counter-intuitively, faster-moving predators lead to lower oscillation periods and eventually to extinction of the predator population. The study does not only provide valuable insights on theoretical spatially explicit predator-prey models with group defense but also comparisons of ecological data with model simulations. © 2020 Elsevier B.V

    A type IV functional response with different shapes in a predator-prey model.

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    Group defense is a phenomenon that occurs in many predator-prey systems. Different functional responses with substantially different properties representing such a mechanism exist. Here, we develop a functional response using timescale separation. A prey-dependent catch rate represents the group defense. The resulting functional response contains a single parameter that controls whether the group defense functional response is saturating or dome-shaped. Based on that, we show that the catch rate must not increase monotonically with increasing prey density to lead to a dome-shaped functional response. We apply bifurcation analysis to show that non-monotonic group defense is usually more successful. However, we also find parameter regions in which a paradox occurs. In this case, higher group defense can give rise to a stable limit cycle, while for lower values, the predator would go extinct. The study does not only provide valuable insight on how to include functional responses representing group defense in mathematical models, but it also clarifies under which circumstances the usage of different functional responses is appropriate
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